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The Star Atop the Tree was the huge “Christmas star” ornament which sat at the top of the giant Christmas tree in the centre of the Christmas Toyland in Madame Tarsa's Toybox. The sentient toys who inhabited the Toyland held the Star to be their realm's greatest treasure.
Unbeknownst to even Millicent, the Star had in fact been placed there by Tarsa to conceal the magical “key” to her resurrection in case of an emergency. Millicent owned a magical tome containing a rhyming riddle which hinted at this, although after having read to them, it took some time for Juliet-178, Arganthone-056 and Jenny Everywhere to figure out that the “distant star” in the poem was that star, rather than a literal celestial body in some other part of the Multiverse. (PROSE: The Time of the Toymaker)